15 years after a global apocalypse, mankind is on the verge of extinction. Civilisation no longer exists, food is scarce and most eke out a living by stealing and killing. One boy clings onto life in the desolate British countryside, where staying away from others has been key to his survival. But this self-imposed isolation comes to an abrupt end when he crosses paths with another group of survivors and faces an enemy far more savage than any of them could imagine.
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A “metal fetishist”, driven mad by the maggots wriggling in the wound he’s made to embed metal into his flesh, runs out into the night and is accidentally run down by a Japanese businessman and his girlfriend. The pair dispose of the corpse in hopes of quietly moving on with their lives. However, the businessman soon finds that he is now plagued by a vicious curse that transforms his flesh into iron.
Sometime in the near future, paraplegic prison inmate Matt Mills meets “The Board” of Satoshi-Telefair Industries, who thank him for joining the Program. Under pressure and with a promise to make him walk again, Mills agrees to volunteer. He is injected with a synthetic nanomaterial and suffocated, only to wake up with restored use of his legs, yet alone in a steel cell in the first spoke of an experimental apparatus – The Wheel. A series of assailants inflict a multitude of damage to Mills’ body and torment his mind, yet the synthetic nanomaterial not only restores his damaged organs, it re-engineers his strength. The experiment might be designed to kill, but Mills is determined to survive – and get back to his daughter before The Board finds her first.
Leela becomes an outlaw when she and a group of ecologically-minded feminists attempt to save an asteroid of primitive life forms and the Violet Dwarf star from being destroyed, while Fry joins a secret society and attempts to stop a mysterious species known as the “Dark Ones” from destroying all life in the universe.